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Most modern crinoids, i.e., the feather stars, are free-moving and lack a stem as adults. Examples of fossil crinoids that have been interpreted as free-swimming include Marsupites, Saccocoma and Uintacrinus.[25] In general, crinoids move to new locations by crawling, using the cirri as legs. Such a movement may be induced in relation to a change in current direction, the need to climb to an elevated perch to feed, or because of an agonistic behaviour by an encountered individual.[26] Crinoids can also swim. They do this by co-ordinated, repeated sequential movements of the arms in three groups. At first the direction of travel is upwards but soon becomes horizontal, travelling at about 7 cm (2.8 in) per second with the oral surface in front. Swimming usually takes place as short bursts of activity lasting up to half a minute, and in the comatulid Florometra serratissima at least, only takes place after mechanical stimulation or as an escape response evoked by a predator.[26] In 2005, a stalked crinoid was recorded pulling itself along the sea floor off the Grand Bahama Island. While it has been known that stalked crinoids could move, before this recording the fastest motion known for a stalked crinoid was 0.6 metres (2 feet) per hour. The 2005 recording showed one of these moving across the seabed at the much faster rate of 4 to 5 cm (1.6 to 2.0 in) per second, or 144 to 180 m (472 to 591 ft) per hoIf one ignores the enigmatic Echmatocrinus of the Burgess Shale, the earliest known unequivocal crinoid groups date back to the Ordovician, 480 million years ago. There are two competing hypotheses pertaining to the origin of the group: the traditional viewpoint holds that crinoids evolved from within the blastozoans (the eocrinoids and their derived descendants, the blastoids and the cystoids), whereas the most popular alternative suggests that the crinoids split early from among the edrioasteroids.[28] The debate is difficult to settle, in part because all three candidate ancestors share many characteristics, including radial symmetry, calcareous plates, and stalked or direct attachment to the substrate
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